At the Great Gallery of Hertford House, home to the Wallace Collection, one of the most significant collections of European fine and decorative arts in the world, the experts being put to the test by Griff Rhys Jones are medieval art historian Dr Janina Ramirez, ceramics specialist historian Lars Tharp, art critic and historian Andrew Graham-Dixon, and historian and director of the Landmark Trust, Anna Keay. The two teams pit their knowledgeable wits against each other in a series of rounds informed by this extraordinary collection.

At the Foundling Museum - home to an extraordinary collection of art and sculpture, often donated by the artists themselves to support Britain's first home for abandoned children - the experts being put to the test by Griff Rhys Jones are historian Professor Kate Williams, medieval art historian Dr Janina Ramirez, author and historian Dan Cruickshank, and cultural historian Dr Gus Casely-Hayford.