Smiley O'Brien from the mirror universe kidnaps Commander Sisko and convines him to impersonate his deceased, mirror universe self in order to persuade the alternate version of his dead wife, Jennifer Sisko, to join the Terran Rebels.

When the covertly assembled Romulan-Cardassian attack fleet enters the Gamma Quadrant on a mission of destruction, Garak is forced to interrogate Odo for information on the Founders. Meanwhile, Sisko goes against orders on a mission to rescue Odo.